Difference between revisions of "Aquilegia formosa"

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[[File:Aquilegia2.jpg|right|275px|Aquilegia formosa]]
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* Scientific Name: ''Aquilegia'' ''formosa''
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* Family: Ranunculaceae
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* Common Names: red columbine, Sitka columbine
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* Codon: AQUFOR
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----
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[[File:AQUFOR1.jpg |thumb|Photo by Bernard Kovalchik, also featured on Main Page]]
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=== Taxonomy ===
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{{Taxobox
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| image =
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| image_caption =Photo by Bernard Kovalchik, also featured on Main Page
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| name = 
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| regnum = [[Plant]]ae
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| subregnum = Viridiplantae
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| phylum = Tracheophyta
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| subphylum= Spermatophytina
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| classis = Magnoliopsida
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| subclassis = Ranunculanae
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| ordo = Ranunculales
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| familia = Ranunculaceae
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| genus = ''Aquilegia'' L.
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| species = '''''Aquilegia formosa''''' Fisch. Ex DC.
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| subspecies =
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| synonyms = *''Aquilegia canadensis'' L. var. ''formosa''
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*''Aquilegia columbiana'' Rydb.
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*''Aquilegia formosa'' Fisch. ex DC. in part  [H&C, KZ99]
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*''Aquilegia formosa'' Fischer ex de Candolle var. ''communis'' B. Boivin
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*''Aquilegia formosa'' Fischer ex de Candolle var. ''megalantha'' B. Boivin
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*''Aquilegia formosa'' Fischer var. ''wawawensis'' (Payson) H. St. John <ref>[http://biology.burke.washington.edu/herbarium/waflora/checklist.php?Taxon=Aquilegia%20formosa%20var.%20formosa Washington Flora Checklist]</ref>
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}}
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<ref>Integrated Taxonomic Information System. Retrieved from https://www.itis.gov/servlet/SingleRpt/SingleRpt</ref>
  
'''''Aquilegia formosa''''', commonly known as the red columbine, western columbine, or the Sitka columbine, is a flowering plant in the Ranunculaceae family. <br><br>
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=== Description ===
'''Synonyms:'''
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Perennial herb growing from a simple to branched woody base with several stems, to 10 dm. tall.<ref name="UW">WTU
*''Aquilegia canadensis L. var. formosa''
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Herbarium, Burke Museum, & University of Washington. Retrieved from http://biology.burke.washington.edu/herbarium/imagecollection/taxon.php?Taxon=Aquilegia%20formosa</ref>  Long petiolate, mostly basal leaves to 5.5 cm long.<ref name="UW" /> Red flowers usually several, nodding; sepals 5, petal-like, short-clawed; petals 5, yellow, red-spurred at base;<ref name=":0">Hitchcock, C. L., Cronquist, A., Giblin, D.,
*''Aquilegia columbiana Rydb.''
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& Legler, B. et al. (2018). ''Flora of the Pacific Northwest: an
*''Aquilegia formosa Fisch. ex DC. in part  [H&C, KZ99]''
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illustrated manual''. Seattle: University of Washington Press. p. 86.</ref> stamens numerous, the inner ones sterile.<ref name="UW" /> Follicles 5.<ref name=":0" />  
*''Aquilegia formosa Fischer ex de Candolle var. communis B. Boivin''
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*''Aquilegia formosa Fischer ex de Candolle var. megalantha B. Boivin''
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*''Aquilegia formosa Fischer var. wawawensis (Payson) H. St. John'' <ref>[http://biology.burke.washington.edu/herbarium/waflora/checklist.php?Taxon=Aquilegia%20formosa%20var.%20formosa Washington Flora Checklist]</ref>
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===Taxonomy===
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*Kingdom Plantae – Plants
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*Subkingdom Tracheobionta – Vascular plants
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*Superdivision Spermatophyta – Seed plants
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*Division Magnoliophyta – Flowering plants
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*Class Magnoliopsida – Dicotyledons
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*Subclass - Magnoliidae
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*Order Ranunculales
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*Family Ranunculaceae – Buttercup family
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*Genus Aquilegia L. – columbine
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*Species Aquilegia formosa Fisch. ex DC. – western columbine
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===Description===
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Densely pubescent perennial from a branched to simple woody base with several stems 1.5-10 dm. tall. Long petiolate, mostly basal leaves 1.5-5.5 cm long.  Pale to deep red flowers usually several, nodding; sepals 5, petal-like, oblong-lanceolate, 1.5-2.5 cm. long; petals 5, yellow, with straight spurs 10-15 mm. long, the blades 2-5 mm. long; stamens numerous, the inner ones sterile and scale-like.  Follicles 5, 2 cm. long, copiously glandular-puberulent. <ref name="UW">[http://biology.burke.washington.edu/herbarium/imagecollection.php WTU Herbarium Profiles]</ref>  
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===Bloom Period===
 
===Bloom Period===
May-August <ref name="UW"/>
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May-August <ref name="UW" />
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[[File:Screen shot 2012-04-30 at 12.30.41 PM.png|right|300px|Distribution of western columbine]]
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===Distribution===
 
===Distribution===
Widely distributed throughout Washington; Alaska south to California, coastal and inland to Alberta and Wyoming. <ref name="UW"/>
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Widely distributed throughout Washington; Alaska south to California, coastal and inland to the Rocky Mountains. <ref name="UW" />
 
===Habitat===
 
===Habitat===
Open woods, lowlands to mid-elevations in the mountains. <ref name="UW"/>
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Open woods, lowlands to mid-elevation. <ref name="UW" />
 
===Uses===
 
===Uses===
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Western columbine is a pollinator food source and thus can be used in site rehabilitation.  It also attracts hummingbirds, browsing and grazing animals.<ref>Goodrich, A. (2012). Guidelines for
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Establishment of Seed Production Sites on Military Installations. Department of Defense Legacy
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Resource Management Program. Retrieved from <nowiki>https://denix.osd.mil/nr/focus-areas/habitat/habitat-restoration/reports/guidelines-of-establishment-of-seed-production-sites-on-military-installations-final-report-february-2012-legacy-10-326/</nowiki></ref>
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Western First Nations have used ''A. formosa'' for many purposes, including the treatment of coughs, stomach aches and diarrhea.<ref>Native
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American Ethnobotany Database. Retrieved from http://naeb.brit.org/uses/search/?string=aquilegia+formosa</ref>
  
 
===Propagation===
 
===Propagation===
Follicles normally dry and split open at maturity. Gently crush dried seed heads to release remaining seeds; cleaned with "office clipper" air-screen. The papery, light pod chaff is easily separated from seed.  Fairly long moist pre-chill improved germination; seeded cones stored for 6 months in a cold walk-in cooler with soil surface kept moist, produced more than 15% germination. Germination tests at Oregon State University seed lab were reported at just 8% with a 3-day prechill treatment. 3 to 5 seeds each were sown into Ray-leach SC-10 super cells filled with Fisons' Sunshine #1 potting mix, amended with 3-month slow-release Osmocote NPK fertilizer and small amounts of Micromax trace elements. Cones were well-watered and placed in to a walk-in cooler at 40 F for 6 months cold-stratification. Cones were moved outdoors to shadehouse in mid-spring to germinate. Initial germination is spotty, and initial growth is quite slow. Seedlings need light but fairly frequent watering to keep soil moist but not soggy.  
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Follicles normally dry and split open at maturity. Gently crush dried seed heads to release remaining seeds; cleaned with "office clipper" air-screen. The papery, light pod chaff is easily separated from seed.  Fairly long moist pre-chill improved germination; seeded cones stored for 6 months in a cold walk-in cooler with soil surface kept moist, produced more than 15% germination. Germination tests at Oregon State University seed lab were reported at just 8% with a 3-day prechill treatment. 3 to 5 seeds each were sown into Ray-leach SC-10 super cells filled with Fisons' Sunshine #1 potting mix, amended with 3-month slow-release Osmocote NPK fertilizer and small amounts of Micromax trace elements. Cones were well-watered and placed in to a walk-in cooler at 40 F for 6 months cold-stratification. Cones were moved outdoors to shadehouse in mid-spring to germinate. Initial germination is spotty, and initial growth is quite slow. Seedlings need light but fairly frequent watering to keep soil moist but not soggy.<ref>Native Plant Network Propagation Protocol Database. Retrieved  from <nowiki>https://npn.rngr.net/renderNPNProtocolDetails?selectedProtocolIds=ranunculaceae-aquilegia-2362</nowiki></ref>
 
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[[File:Aquilegia formosa.jpg|300px|thumb|''Aquilegia formosa'', photo by Lisa Hintz]]
 
===Seed===
 
===Seed===
'''Abbreviation:''' AQFO
 
  
 
'''Seed sample from:''' 2011
 
'''Seed sample from:''' 2011
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'''Longitudinal Cross Section:''' obovate [[File:AQFO long.png]]
 
'''Longitudinal Cross Section:''' obovate [[File:AQFO long.png]]
  
[[File:Aquilegia formosa.jpg|300px|thumb|left|''Aquilegia formosa'']]
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[[File:Aquilegia formosa single.jpg|300px|thumb|right|''Aquilegia formosa'']]
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[[File:Aquilegia formosa single.jpg|300px|thumb|''Aquilegia formosa,'' photo by Lisa Hintz]]
  
  
  
 
{{Basics}}
 
{{Basics}}
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===Photo Gallery===
 
===Photo Gallery===
 
<gallery>
 
<gallery>
File:Aquilegia2.jpg
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File: AQUFOR2.jpg| Photo by Donovan Tracy, flowersofrainier.com
File:Aquilegia formosa.jpg
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File:AQUFOR3.jpg| seedling, courtesy of CNLM
File:Aquilegia formosa single.jpg
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File:AQUFOR4.jpg| seedling, courtesy of CNLM
 
</gallery>
 
</gallery>
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===References===
 
===References===
<References/>
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<references />

Latest revision as of 14:20, 24 June 2021

  • Scientific Name: Aquilegia formosa
  • Family: Ranunculaceae
  • Common Names: red columbine, Sitka columbine
  • Codon: AQUFOR

Photo by Bernard Kovalchik, also featured on Main Page

Taxonomy

Scientific classification
Kingdom: Plantae
Subkingdom: Viridiplantae
Phylum: Tracheophyta
Subphylum: Spermatophytina
Class: Magnoliopsida
Subclass: Ranunculanae
Order: Ranunculales
Family: Ranunculaceae
Genus: Aquilegia L.
Species: Aquilegia formosa Fisch. Ex DC.
Synonyms
  • Aquilegia canadensis L. var. formosa
  • Aquilegia columbiana Rydb.
  • Aquilegia formosa Fisch. ex DC. in part [H&C, KZ99]
  • Aquilegia formosa Fischer ex de Candolle var. communis B. Boivin
  • Aquilegia formosa Fischer ex de Candolle var. megalantha B. Boivin
  • Aquilegia formosa Fischer var. wawawensis (Payson) H. St. John [1]

[2]

Description

Perennial herb growing from a simple to branched woody base with several stems, to 10 dm. tall.[3] Long petiolate, mostly basal leaves to 5.5 cm long.[3] Red flowers usually several, nodding; sepals 5, petal-like, short-clawed; petals 5, yellow, red-spurred at base;[4] stamens numerous, the inner ones sterile.[3] Follicles 5.[4]

Bloom Period

May-August [3]

Distribution of western columbine

Distribution

Widely distributed throughout Washington; Alaska south to California, coastal and inland to the Rocky Mountains. [3]

Habitat

Open woods, lowlands to mid-elevation. [3]

Uses

Western columbine is a pollinator food source and thus can be used in site rehabilitation. It also attracts hummingbirds, browsing and grazing animals.[5] Western First Nations have used A. formosa for many purposes, including the treatment of coughs, stomach aches and diarrhea.[6]

Propagation

Follicles normally dry and split open at maturity. Gently crush dried seed heads to release remaining seeds; cleaned with "office clipper" air-screen. The papery, light pod chaff is easily separated from seed. Fairly long moist pre-chill improved germination; seeded cones stored for 6 months in a cold walk-in cooler with soil surface kept moist, produced more than 15% germination. Germination tests at Oregon State University seed lab were reported at just 8% with a 3-day prechill treatment. 3 to 5 seeds each were sown into Ray-leach SC-10 super cells filled with Fisons' Sunshine #1 potting mix, amended with 3-month slow-release Osmocote NPK fertilizer and small amounts of Micromax trace elements. Cones were well-watered and placed in to a walk-in cooler at 40 F for 6 months cold-stratification. Cones were moved outdoors to shadehouse in mid-spring to germinate. Initial germination is spotty, and initial growth is quite slow. Seedlings need light but fairly frequent watering to keep soil moist but not soggy.[7]

Aquilegia formosa, photo by Lisa Hintz

Seed

Seed sample from: 2011

Average Measurement: 2.5 x 1.4 x 1.2

Measurement Range: L: 2.5 - 2.75, W: 1.2 - 1.5, D: 1 - 1.5

Features

Shape: Seeds tapering at hilum end, broadening at opposite side.

Color: Hilum is white, seed is black.

Surface: At least one distinct ridge runs from the hilum to the opposite side. End opposite hilum is sometimes puckered or having concave pits. Seed is glossy and lightly textured with very small bumps.

Latitudinal Cross Section: elliptical AQFO lat.png

Longitudinal Cross Section: obovate AQFO long.png


Aquilegia formosa, photo by Lisa Hintz


Basic Explanations and Assumptions:

The dimensions for the seeds are length x width x depth. The location of the hilum is used as the base of the seed, and the length is measured from hilum to the opposite apex. Where a style is present, the length is measured from the hilum to the bottom of the style. Width is measured at a right angle to the length at the widest part. Depth is measured at a right angle to the intersection of height and width lines.

Measurements included are the mean average for each measurement of ten separate seeds.

All measurements in millimeters unless otherwise noted.

Photo Gallery

References

  1. Washington Flora Checklist
  2. Integrated Taxonomic Information System. Retrieved from https://www.itis.gov/servlet/SingleRpt/SingleRpt
  3. 3.0 3.1 3.2 3.3 3.4 3.5 WTU Herbarium, Burke Museum, & University of Washington. Retrieved from http://biology.burke.washington.edu/herbarium/imagecollection/taxon.php?Taxon=Aquilegia%20formosa
  4. 4.0 4.1 Hitchcock, C. L., Cronquist, A., Giblin, D., & Legler, B. et al. (2018). Flora of the Pacific Northwest: an illustrated manual. Seattle: University of Washington Press. p. 86.
  5. Goodrich, A. (2012). Guidelines for Establishment of Seed Production Sites on Military Installations. Department of Defense Legacy Resource Management Program. Retrieved from https://denix.osd.mil/nr/focus-areas/habitat/habitat-restoration/reports/guidelines-of-establishment-of-seed-production-sites-on-military-installations-final-report-february-2012-legacy-10-326/
  6. Native American Ethnobotany Database. Retrieved from http://naeb.brit.org/uses/search/?string=aquilegia+formosa
  7. Native Plant Network Propagation Protocol Database. Retrieved from https://npn.rngr.net/renderNPNProtocolDetails?selectedProtocolIds=ranunculaceae-aquilegia-2362