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''Dichanthelium acuminatum (Sw.) Gould & Clark'', also called tapered rosette grass, Western panicum, Western witchgrass, or Western panicgrass, is a perennial native plant of the Puget Prairie ecosystem in the Poaceae family. Other names: ''Panicum occidentale''.
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''Dodecatheon pulchellum (Raf.) Merr.'', also called Shooting Star, Dark-throated shooting star, Sticky shooting star, Cusick's shooting star, Few-flowered shooting star, Western shooting star. This native perennial Puget prairie forb/herb belongs to Primulaceae, the primrose family. Other names: (English, First People's, etc.)
 
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Line 9: Line 8:
 
*Superdivision Spermatophyta – Seed plants <br>
 
*Superdivision Spermatophyta – Seed plants <br>
 
*Division Magnoliophyta – Flowering plants <br>
 
*Division Magnoliophyta – Flowering plants <br>
*Class Liliopsida Monocotyledons <br>
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*Class Magnoliopsida Dicotyledons <br>
*Subclass Commelinidae <br>
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*Subclass Dilleniidae <br>
*Order Cyperales <br>
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*Order Primulales <br>
*Family Poaceae Grass family <br>
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*Family Primulaceae Primrose family <br>
*Genus Dichanthelium (Hitchc. & Chase) Gould yarrow <br>
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*Genus Dodecatheon L. – shootingstar <br>
*Species Dichanthelium acuminatum (Sw.) Gould & C.A. Clark tapered rosette grass
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*Species Dodecatheon pulchellum (Raf.) Merr. – darkthroat shootingstar
  
 
==Description==
 
==Description==
  
Tufted perennial; stems yellowish-green, leafy towards base, spreading, usually velvety-hairy, 15-40 cm tall. Leaves flat, firm, erect to ascending, hairy, 5-10 mm wide, no auricles; ligules 3-4 mm long, consisting of long hairs. Inflorescence a panicle, open, 3-9 cm long; spikelets up to 2 mm long, short-hairy, two flowered, the lower flower sterile; glumes unequal in size; fertile lemmas hardened (Pojar and MacKinnon, 1994)
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General: Scapose, perennial herbs from short rhizomes, glabrous to glandular-pubescent throughout, the flowering stems 5-40 cm. tall.
 +
Leaves: Leaves basal, the blades oblong-lanceolate to oblanceolate, 2-15 cm. long, entire to denticulate, narrowed gradually to winged petioles nearly as long.
 +
Flowers: Flowers 1-25 in an umbel, terminal on the scape, 5-parted; calyx purple-flecked, the lobes 3-5 mm. long; corolla 10-20 mm. long, the lobes purplish-lavender, the tube yellowish with a purple line at the base; stamens clustered around the style, opposite the petals, the filaments united in a yellow, orange or purple tube 1.5-3 mm. long; anthers yellowish to reddish-purple, 4-7 mm. long; style single, barely exceeding the stamens, the stigma slightly enlarged
 +
Fruit: Capsule ovoid-cylindric, 5-15 mm. long, glabrous to glandular hairy, opening by valves to the tip.
  
[[Image:diac2_001_lvp.jpg|thumb|right|Robert H. Mohlenbrock @ USDA-NRCS PLANTS Database / USDA SCS. 1989. Midwest wetland flora: Field office illustrated guide to plant species. Midwest National Technical Center, Lincoln.]]
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[[Image:dopu_001_pvp.jpg|thumb|right|Mr. & Mrs. Robert G. Young. USDA NRCS. 1992. ''Western wetland flora: Field office guide to plant species''. West Region, Sacramento. Courtesy of USDA NRCS Wetland Science Institute.]]
  
 
==Bloom Period==
 
==Bloom Period==
June-September
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Late Spring
 
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Dichantheliums are characterized by two distinct blooming periods. The conspicuous primary flowering heads are terminal to the culms and are produced in late spring and early summer. Secondary flowering heads are produced from the leaf axils begininning in mid-summer and continuing into early autumn. The primary flowering heads usually have a lower seedset than the secondary ones, which have flowers that remain closed and are self pollinated. However, seeds produced by the primary flowers appear to germinate more readily than seeds from the secondary flowers.
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(nativeplantnetwork.org)
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==Distribution==
 
==Distribution==
Southern British Columbia south along the coast to California; east of the Cascades mainly along water courses or around springs in the mountains, east to Montana and Wyoming.
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Alaska to Mexico, east to Pennsylvania; found throughout the Pacific Northwest. It ranges from Alaska to Manitoba south to California, Colorado, Nebraska, and Arizona.  
  
 
==Habitat==
 
==Habitat==
Rocky or sandy river banks or lake margins to open woods, marshy areas or dry prairies, from sea level to high elevation in the mountains. Moist, sandy ground (lakeshores, beaches, stream banks), rocky and gravelly areas in bogs, meadows and open forests at low to mid elevations (Pojar and MacKinnon, 1994).
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''Dodecatheon pulchellum'' occurs in perennially moist to wet soils of meadows and along streams from mid elevations in the mountains to the alpine. Its ecological setting ranges from Coastal prairies to inland saline swamps and shrub-steppe, sea level to timberline.
  
 
==Uses==
 
==Uses==
Landscaping: Panic grass is good for a low-traffic ground cover. Useful ground cover in controlled moist area. Attractive fall/winter color. Excellent in poorly-drained locations (S. Bastin, pers. comm.).
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Low palatability for browsing and grazing animals.
  
 
==Propagation==
 
==Propagation==
Seed germinates readily with day temperatures of 20º C and night temperatures of 5-10º C Do not cover seeds; simply lightly press into the soil. Requires continuous moisture for germination. Good results will fall sowing - 90% germination by mid-April (S. Bastin, pers. comm.).
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Citations:
 +
Evans, Jeff; Hosokawa, Joy.; Wick, Dale. 2008. Propagation protocol for production of container Dodecatheon pulchellum (Raf.) Merr. plants (172 ml containers); USDI NPS - Glacier National Park, West Glacier, Montana. In: Native Plant Network. URL: http://www.nativeplantnetwork.org (accessed 8 March 2010). Moscow (ID): University of Idaho, College of Natural Resources, Forest Research Nursery
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Skinner, David M. 2006. Propagation protocol for production of container Dodecatheon pulchellum (Raf.) Merr. plants (10 cu. in.); USDA NRCS - Pullman Plant Materials Center, Pullman, Washington. In: Native Plant Network. URL: http://www.nativeplantnetwork.org (accessed 8 March 2010). Moscow (ID): University of Idaho, College of Natural Resources, Forest Research Nursery.
 +
 
 +
Seed Storage
 +
Storage Requirements: Refrigerator.
 +
Seed Storage Longevity:
 +
Germination: 65% (Evans, Hosokawa & Wick).
 +
 
 +
Clean seed is stored at 40 degrees Fahrenheit and 40% relative humidity (Skinner).
 +
 
 +
Nursery Conditions
 +
Infrastructure Requirements:  Outdoor nursery growing facility.
 +
Nutrition Program:
 +
Pest Control Practices:
 +
Special Growing Medium Requirements:  Growing medium used is 6:1:1 milled spaghnum peat, perlite, and vermiculite with Osmocote controlled release fertilizer (13N:13P2O5:13K2O; 8 to 9 month release rate at 21C) and Micromax fertilizer (12%S, 0.1%B, 0.5%Cu, 12%Fe, 2.5%Mn, 0.05%Mo, 1%Zn) at the rate of 1 gram of Osmocote and 0.20 gram of Micromax per 172 ml container (Evans, Hosokawa & Wick).
 +
Cross-Pollination Concerns:
 +
Time to Grow: 8 months (Evans, Hosokawa & Wick). 3 years (Skinner).
 +
Timeline:  Sow Nov,Germinate May, Harvest July (Evans et al).
 +
 
 +
Production Phase 1- Fruit/Seed Dormancy and Pre-Treatment
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Length of Dormancy:
 +
Type of dormancy:  Physiological dormancy (Evans, Hosokawa & Wick)
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Pretreatment Type:  Outdoor cold, moist stratification (Evans, Hosokawa & Wick). 
 +
Pretreatment Length: 5 months (Evans, Hosokawa & Wick)
 +
 
 +
Extended cold, moist stratification is needed. Cool spring temperatures may also be necessary. In trials at the PMC, no germination occurred without stratification and no seed germinated after 30 days cold, moist stratification. Seed planted in containers in November and left outside began emerging in late March and reached 65% by late April.
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Seed of Dodecatheon meadia requires 2 months of cool, moist stratification (Greene & Curtis 1950) (Skinner).
 +
 
 +
Production Phase 2- Sowing and Germination
 +
Sowing Methodologies:  In early November seed is sown in 10 cu. in. Ray Leach Super cells filled with Sunshine #4 and covered lightly. A thin layer of pea gravel is applied to prevent seeds from floating. Containers are watered deeply and placed outside (Skinner).
 +
Containers are filled and sown in late fall and irrigated thoroughly prior to winter stratification (Evans et al).
 +
 
 +
Recommended Sow Rate:
 +
Germination Length:(time from sowing to emergence of radicle)
  
Conditioned seed is planted into round cell greenhouse flat liners with 38 cells per flat that have been filled with coarse processed bark and composted pine bark growing medium. Seed is surface sown at a rate of 3-5 seeds per cell and lightly covered with starter sized, 1/16" - 1/8" diameter, granite poultry grit to combat damping off diseases. Prepared flats are lightly hand watered to slightly moisten the growing medium.  
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Production Phase 3- Establishment
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Establishment Methodology:  Containers remain outside. They are watered only during dry spells. Plants begin emerging in late March. Germination occurs over a 3-4 week period (Skinner).
 +
Seedlings germinate in spring under fluctuating outdoor temperatures and are grown under full sun exposure. Seedlings are irrigated with Rainbird automatic irrigation system in early morning until containers are thoroughly leached.  Germination occurs at fluctuating temperatures during May. Medium is kept slightly moist during germination. (Evans et al).
 +
Establishment Length:  4 weeks (Skinner) (Evans et al).
  
Stratified seed is placed in a greenhouse maintained under natural lighting and at a minimum temperature of 70 degrees Fahrenheit. Soil moisture is maintained during germination by an automatic overhead watering system set to cycle for 20 seconds every thirty minutes during daylight hours.
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Production Phase 4-Active Growth
Germination typically occurs 7 - 10 days after placement in the greenhouse.
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Active Growth Methodology:  Plants are watered as needed while outside and fertilized once a week with a water soluble, complete fertilizer. They are moved to the lath house in June. The plants develop small rosettes with several true leaves. They will not go dormant with summer temperatures but cease growing (Skinner).
After germination, seedlings are maintained in a greenhouse environment 2-4 months to promote development of a plug with at least 6 inches of top growth and a dense, fibrous root system suitable for mechanical transplanting. Watering is reduced to overhead hand watering once daily. Seedlings receive a water soluble complete fertilizer bi-weekly until hardening.  
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Root and shoot development occurs rapidly following germination. 4 to 6 true leaves were evident 3 weeks after germination. Seedlings were root tight 4 weeks after germination (Evans et al).
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Active Growth Length:  2 months (Skinner). 12 weeks (Evans et al).
  
Hardening Phase 1 – 2 weeks.
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Production Phase 5- Finishing
 +
Finishing/Hardening Methodology: Fertilizer is withheld in September and water decreased to harden the plants for winter. Since the plants are grown outside, additional hardening is not needed (Skinner).
 +
Plants are fertilized with 10-20-20 liquid NPK at 200 ppm during August and September. Irrigation is gradually reduced in September and October. Plants were given one final irrigation prior to winterization (Evans et al).
 +
Finishing/Hardening Length: 4 weeks (Evans et al).
 +
Overwinter live plant storage:  Overwinter in outdoor nursery under insulating foam cover and snow (Evans et al).
 +
Plants are stored in the lath house over winter. They should be afforded some protection from extreme cold temperatures. Mulch or foam sheets provide sufficient protection. The protection should be removed in late winter or early spring as temperatures begin to rise (Skinner).
  
(nativeplantnetwork.org)
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Production Phase 6- Outplanting Indications:
 +
Because the plants begin growing early in the spring, late fall outplanting is preferred where soils are generally too muddy in February and early March. However, the tapered plug from the container has a tendency to frost heave and leave the upper part of the plug exposed. Fall transplants should be checked in early spring and exposure of the plug corrected.
 +
Transplanting is done in late October by using an electric drill and portable generator to drill 1.5 inch diameter holes at the planting site.
 +
Survival in a seed increase planting without competing vegetation was 60%. A few plants flowered the following spring (Skinner).
  
 +
Additional Comments:
 +
Dodecatheon species, such as D. jeffreyi, D. conjugens, D. pulchellum are reported to germinate best when subjected to a cold, moist stratification for at least 90 days (Evans et al).
 +
Plants should not be dug up from stands in the wild. Wild grown plants rarely survive the transplanting process (Parish 1996, Kruckeberg 1996).
  
 
===Seed===
 
===Seed===
[[File:Dichanthelium acuminatum.jpg|300px|thumb|right|''Dichanthelium acuminatum'']]
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[[File:Dodecatheon pulchellum.jpg|300px|thumb|right|''Dodecatheon pulchellum'']]
'''Abbreviation:''' DIAC
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'''Abbreviation:''' DOPU
  
'''Seed sample from:''' 2011
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'''Seed sample from:''' 2010
  
 
'''Average Measurement:''' 1.6 x 1 x 0.8
 
'''Average Measurement:''' 1.6 x 1 x 0.8

Revision as of 12:57, 7 May 2012

Dodecatheon pulchellum (Raf.) Merr., also called Shooting Star, Dark-throated shooting star, Sticky shooting star, Cusick's shooting star, Few-flowered shooting star, Western shooting star. This native perennial Puget prairie forb/herb belongs to Primulaceae, the primrose family. Other names: (English, First People's, etc.)


Taxonomy

  • Kingdom Plantae – Plants
  • Subkingdom Tracheobionta – Vascular plants
  • Superdivision Spermatophyta – Seed plants
  • Division Magnoliophyta – Flowering plants
  • Class Magnoliopsida – Dicotyledons
  • Subclass Dilleniidae
  • Order Primulales
  • Family Primulaceae – Primrose family
  • Genus Dodecatheon L. – shootingstar
  • Species Dodecatheon pulchellum (Raf.) Merr. – darkthroat shootingstar

Description

General: Scapose, perennial herbs from short rhizomes, glabrous to glandular-pubescent throughout, the flowering stems 5-40 cm. tall. Leaves: Leaves basal, the blades oblong-lanceolate to oblanceolate, 2-15 cm. long, entire to denticulate, narrowed gradually to winged petioles nearly as long. Flowers: Flowers 1-25 in an umbel, terminal on the scape, 5-parted; calyx purple-flecked, the lobes 3-5 mm. long; corolla 10-20 mm. long, the lobes purplish-lavender, the tube yellowish with a purple line at the base; stamens clustered around the style, opposite the petals, the filaments united in a yellow, orange or purple tube 1.5-3 mm. long; anthers yellowish to reddish-purple, 4-7 mm. long; style single, barely exceeding the stamens, the stigma slightly enlarged Fruit: Capsule ovoid-cylindric, 5-15 mm. long, glabrous to glandular hairy, opening by valves to the tip.

Mr. & Mrs. Robert G. Young. USDA NRCS. 1992. Western wetland flora: Field office guide to plant species. West Region, Sacramento. Courtesy of USDA NRCS Wetland Science Institute.

Bloom Period

Late Spring

Distribution

Alaska to Mexico, east to Pennsylvania; found throughout the Pacific Northwest. It ranges from Alaska to Manitoba south to California, Colorado, Nebraska, and Arizona.

Habitat

Dodecatheon pulchellum occurs in perennially moist to wet soils of meadows and along streams from mid elevations in the mountains to the alpine. Its ecological setting ranges from Coastal prairies to inland saline swamps and shrub-steppe, sea level to timberline.

Uses

Low palatability for browsing and grazing animals.

Propagation

Citations: Evans, Jeff; Hosokawa, Joy.; Wick, Dale. 2008. Propagation protocol for production of container Dodecatheon pulchellum (Raf.) Merr. plants (172 ml containers); USDI NPS - Glacier National Park, West Glacier, Montana. In: Native Plant Network. URL: http://www.nativeplantnetwork.org (accessed 8 March 2010). Moscow (ID): University of Idaho, College of Natural Resources, Forest Research Nursery Skinner, David M. 2006. Propagation protocol for production of container Dodecatheon pulchellum (Raf.) Merr. plants (10 cu. in.); USDA NRCS - Pullman Plant Materials Center, Pullman, Washington. In: Native Plant Network. URL: http://www.nativeplantnetwork.org (accessed 8 March 2010). Moscow (ID): University of Idaho, College of Natural Resources, Forest Research Nursery.

Seed Storage Storage Requirements: Refrigerator. Seed Storage Longevity: Germination: 65% (Evans, Hosokawa & Wick).

Clean seed is stored at 40 degrees Fahrenheit and 40% relative humidity (Skinner).

Nursery Conditions Infrastructure Requirements: Outdoor nursery growing facility. Nutrition Program: Pest Control Practices: Special Growing Medium Requirements: Growing medium used is 6:1:1 milled spaghnum peat, perlite, and vermiculite with Osmocote controlled release fertilizer (13N:13P2O5:13K2O; 8 to 9 month release rate at 21C) and Micromax fertilizer (12%S, 0.1%B, 0.5%Cu, 12%Fe, 2.5%Mn, 0.05%Mo, 1%Zn) at the rate of 1 gram of Osmocote and 0.20 gram of Micromax per 172 ml container (Evans, Hosokawa & Wick). Cross-Pollination Concerns: Time to Grow: 8 months (Evans, Hosokawa & Wick). 3 years (Skinner). Timeline: Sow Nov,Germinate May, Harvest July (Evans et al).

Production Phase 1- Fruit/Seed Dormancy and Pre-Treatment Length of Dormancy: Type of dormancy: Physiological dormancy (Evans, Hosokawa & Wick) Pretreatment Type: Outdoor cold, moist stratification (Evans, Hosokawa & Wick). Pretreatment Length: 5 months (Evans, Hosokawa & Wick)

Extended cold, moist stratification is needed. Cool spring temperatures may also be necessary. In trials at the PMC, no germination occurred without stratification and no seed germinated after 30 days cold, moist stratification. Seed planted in containers in November and left outside began emerging in late March and reached 65% by late April. Seed of Dodecatheon meadia requires 2 months of cool, moist stratification (Greene & Curtis 1950) (Skinner).

Production Phase 2- Sowing and Germination Sowing Methodologies: In early November seed is sown in 10 cu. in. Ray Leach Super cells filled with Sunshine #4 and covered lightly. A thin layer of pea gravel is applied to prevent seeds from floating. Containers are watered deeply and placed outside (Skinner). Containers are filled and sown in late fall and irrigated thoroughly prior to winter stratification (Evans et al).

Recommended Sow Rate: Germination Length:(time from sowing to emergence of radicle)

Production Phase 3- Establishment Establishment Methodology: Containers remain outside. They are watered only during dry spells. Plants begin emerging in late March. Germination occurs over a 3-4 week period (Skinner). Seedlings germinate in spring under fluctuating outdoor temperatures and are grown under full sun exposure. Seedlings are irrigated with Rainbird automatic irrigation system in early morning until containers are thoroughly leached. Germination occurs at fluctuating temperatures during May. Medium is kept slightly moist during germination. (Evans et al). Establishment Length: 4 weeks (Skinner) (Evans et al).

Production Phase 4-Active Growth Active Growth Methodology: Plants are watered as needed while outside and fertilized once a week with a water soluble, complete fertilizer. They are moved to the lath house in June. The plants develop small rosettes with several true leaves. They will not go dormant with summer temperatures but cease growing (Skinner). Root and shoot development occurs rapidly following germination. 4 to 6 true leaves were evident 3 weeks after germination. Seedlings were root tight 4 weeks after germination (Evans et al). Active Growth Length: 2 months (Skinner). 12 weeks (Evans et al).

Production Phase 5- Finishing Finishing/Hardening Methodology: Fertilizer is withheld in September and water decreased to harden the plants for winter. Since the plants are grown outside, additional hardening is not needed (Skinner). Plants are fertilized with 10-20-20 liquid NPK at 200 ppm during August and September. Irrigation is gradually reduced in September and October. Plants were given one final irrigation prior to winterization (Evans et al). Finishing/Hardening Length: 4 weeks (Evans et al). Overwinter live plant storage: Overwinter in outdoor nursery under insulating foam cover and snow (Evans et al). Plants are stored in the lath house over winter. They should be afforded some protection from extreme cold temperatures. Mulch or foam sheets provide sufficient protection. The protection should be removed in late winter or early spring as temperatures begin to rise (Skinner).

Production Phase 6- Outplanting Indications: Because the plants begin growing early in the spring, late fall outplanting is preferred where soils are generally too muddy in February and early March. However, the tapered plug from the container has a tendency to frost heave and leave the upper part of the plug exposed. Fall transplants should be checked in early spring and exposure of the plug corrected. Transplanting is done in late October by using an electric drill and portable generator to drill 1.5 inch diameter holes at the planting site. Survival in a seed increase planting without competing vegetation was 60%. A few plants flowered the following spring (Skinner).

Additional Comments: Dodecatheon species, such as D. jeffreyi, D. conjugens, D. pulchellum are reported to germinate best when subjected to a cold, moist stratification for at least 90 days (Evans et al). Plants should not be dug up from stands in the wild. Wild grown plants rarely survive the transplanting process (Parish 1996, Kruckeberg 1996).

Seed

Dodecatheon pulchellum

Abbreviation: DOPU

Seed sample from: 2010

Average Measurement: 1.6 x 1 x 0.8

Measurement Range: L: 1.5 x 1.9, W: no variation in sample set (all seeds measured 1 mm) D: 0.7 - 0.9

Features

Shape: One side of seed has two grooves running from hilium to opposite apex that form a football shaped opening.

Color: Seeds in green and purple husk that is coarsely hairy. Inner seed is off-white with a white, slightly puckered hilium.

Surface: Seeds are glossy and smooth. Some are very finely striped longitudinally.

Latitudinal Cross Section: elliptical DIAC lat.png
Longitudinal Cross Section: elliptical DIAC long.png

References

  • Dichanthelium acuminatum - Wikipedia: The Free Encyclopedia. Wikimedia Foundation Inc. [1]. Retrieved 04/12/2012.


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